Saltwater Forest is a Dacrydium cupressinum-dominated lowland forest covering 9000 ha in south Westland, South Island, New Zealand. Four thousand hectares is managed for sustainable production of indigenous timber. The aim of this study was to provide an integrated analysis of soils, soil-landform relationships, and soil-vegetation relationships at broad and detailed scales. The broad scale understandings provide a framework in which existing or future studies can be placed and the detailed studies elucidate sources of soil and forest variability.
Glacial landforms dominate. They include late Pleistocene lateral, terminal and ablation moraines, and outwash aggradation and degradation terraces. Deposits and landforms from six glacial advances have been recognised ranging from latest Last (Otira) Glaciation to Penultimate (Waimea) Glaciation. The absolute ages of landforms were established by analysis of the thickness and soil stratigraphy of loess coverbeds, augmented with radiocarbon dating and phytolith and pollen analysis.
In the prevailing high rainfall of Westland soil formation is rapid. The rate of loess accretion in Saltwater
Forest (ca. 30 mm ka⁻¹) has been low enough that soil formation and loess accretion took place contemporaneously. Soils formed in this manner are known as upbuilding soils. The significant difference between upbuilding pedogenesis and pedogenesis in a topdown sense into an existing sediment body is that each subsoil increment of an upbuilding soil has experienced processes of all horizons above. In Saltwater Forest subsoils of upbuilding soils are strongly altered because they have experienced the extremely acid environment of the soil surface at some earlier time. Some soil chronosequence studies in Westland have included upbuilding soils formed in loess as the older members of the sequence. Rates and types of processes inferred from these soils should be reviewed because upbuilding is a different pedogenic pathway to topdown pedogenesis.
Landform age and morphology were used as a primary stratification for a study of the soil pattern and nature of soil variability in the 4000 ha production area of Saltwater Forest. The age of landforms (> 14 ka) and rapid soil formation mean that soils are uniformly strongly weathered and leached. Soils include Humic Organic Soils, Perch-gley Podzols, Acid Gley Soils, Allophanic Brown Soils, and Orthic or Pan Podzols. The major influence on the nature of soils is site hydrology which is determined by macroscale features of landforms (slope, relief, drainage density), mesoscale effects related to position on landforms, and microscale influences determined by microtopography and individual tree effects. Much of the soil variability arises at microscales so that it is not possible to map areas of uniform soils at practical map scales. The distribution of soil variability across spatial scales, in relation to the intensity of forest management, dictates that it is most appropriate to map soil complexes with boundaries coinciding with landforms.
Disturbance of canopy trees is an important agent in forest dynamics. The frequency of forest disturbance in the production area of Saltwater Forest varies in a systematic way among landforms in accord with changes in abundance of different soils. The frequency of forest turnover is highest on landforms with the greatest abundance of extremely poorly-drained Organic Soils. As the abundance of better-drained soils increases the frequency of forest turnover declines. Changes in turnover frequency are reflected in the mean size and density of canopy trees (Dacrydium cupressinum) among landforms. Terrace and ablation moraine landforms with the greatest abundance of extremely poorly-drained soils have on average the smallest trees growing most densely. The steep lateral moraines, characterised by well drained soils, have fewer, larger trees. The changes manifested at the landform scale are an integration of processes operating over much shorter range as a result of short-range soil variability. The systematic changes in forest structure and turnover frequency among landforms and soils have important implications for sustainable forest management.
This report reviews the literature on regeneration requirements of main canopy
tree species in Westland. Forests managed for production purposes have to be
harvested in an ecologically sustainable way; to maintain their natural character, harvesting should facilitate regeneration of target species and ensure that their recruitment is in proportion to the extent of extraction. The reasons for species establishing at any point in time are unclear; however, they are probably related to the availability of suitable microsites for establishment, the size of the canopy openings formed by disturbance, and whether or not seeds are available at or around the time of the disturbance. Age structures from
throughout Westland show that extensive, similar-aged, post-earthquake cohorts of trees are a feature of the region. This suggests that infrequent, massive earthquakes are the dominant coarse-scale disturbance agent, triggering episodes of major erosion and sedimentation and leaving a strong imprint in the forest structure. In other forests, flooding and catastrophic
windthrow are major forms of disturbance. The findings suggest that, in general, large disturbances are required for conifer regeneration. This has implications for any sustained yield management of these forests if conifers are to remain an important component. Any harvesting should recognise the importance for tree establishment of: forest floor microsites, such as fallen logs
and tree tip-up mounds; and the variable way in which canopy gaps are formed. Harvesting should maintain the 'patchy' nature of the natural forest—large patches of dense conifers interspersed with more heterogeneous patches of mixed species.This is a client report commissioned by West Coast Conservancy and funded from the Unprogrammed Science Advice fund.
Tree mortality is a fundamental process governing forest dynamics, but understanding tree mortality patterns is challenging
because large, long-term datasets are required. Describing size-specific mortality patterns can be especially difficult, due to
few trees in larger size classes. We used permanent plot data from Nothofagus solandri var. cliffortioides (mountain beech)
forest on the eastern slopes of the Southern Alps, New Zealand, where the fates of trees on 250 plots of 0.04 ha were
followed, to examine: (1) patterns of size-specific mortality over three consecutive periods spanning 30 years, each
characterised by different disturbance, and (2) the strength and direction of neighbourhood crowding effects on sizespecific
mortality rates. We found that the size-specific mortality function was U-shaped over the 30-year period as well as
within two shorter periods characterised by small-scale pinhole beetle and windthrow disturbance. During a third period,
characterised by earthquake disturbance, tree mortality was less size dependent. Small trees (,20 cm in diameter) were
more likely to die, in all three periods, if surrounded by a high basal area of larger neighbours, suggesting that sizeasymmetric
competition for light was a major cause of mortality. In contrast, large trees ($20 cm in diameter) were more
likely to die in the first period if they had few neighbours, indicating that positive crowding effects were sometimes
important for survival of large trees. Overall our results suggest that temporal variability in size-specific mortality patterns,
and positive interactions between large trees, may sometimes need to be incorporated into models of forest dynamics.
Knowledge of past climate variability is essential for understanding present and future climate trends. This study used Halocarpus biformis (pink pine) ring-width chronologies to investigate palaeotemperature history in Westland, New Zealand. The ensuing reconstruction is among the longest palaeoseries produced for New Zealand to date. It is in good agreement with other tree-ring-based records, and with instrumental (both local and hemispheric) data.
Thirteen pink pine chronologies were developed. Ring-width measurements were detrended using the Regional Curve Standardisation method to retain as much low-frequency variance as possible. Crossdating revealed the existence of a strong common signal among trees. Inter-site comparison indicated that a common control mechanism affected tree growth not only within sites, but also across sites.
To determine whether climate was the main factor that controlled the growth of pink pine in Westland, correlation and response function analyses were employed. Temperature, precipitation and the Southern Oscillation Index were tested for their relationship with tree growth. Mean monthly temperature was identified as the primary growth-limiting factor. Chronologies were positively correlated with temperature over an extended period (5-17 months), and climate response modelling showed that temperature explained 11-60% variance in the tree-ring data. The highest and most stable correlations occurred between tree growth and summer (January-March) temperatures.
Tree-ring data from the six sites that contained the strongest temperature signal were combined, and the Westland Regional Chronology (WRC) was developed. The WRC was then used to reconstruct January-March temperatures back to A.D. 1480. The calibration model explained 43% of the variance in temperature, and all calibration and verification tests were passed at high levels of significance. The reconstruction showed that temperatures in Westland have been following a positive trend over the last 520 years. The coolest 25-year period was 1542-1566, while temperatures reached their maximum in 1966-1990. Spectral analysis of the Westland palaeotemperature record revealed cycles at periods of about 3, 5-6, 11, 14, 22, 45 and 125 years.
This study also confirmed that climate response is species-dependent. A separate exercise, which compared two species from the same site, demonstrated that while pink pine's growth was mainly influenced by summer temperatures, Libocedrus bidwillii was affected by conditions at the beginning of the growing season. However, the temperature signal in Westland's Libocedrus bidwillii was insufficient to produce a reliable reconstruction. It might be because the climate signal in this species was obscured by disturbances, as was shown in the final section of this project. Frequent growth releases and suppressions implied that Libocedrus bidwillii integrated both major (Alpine Fault earthquakes) and minor (windthrow) disturbances in its ring widths. Pink pine, on the other hand, was not sensitive to disturbance, and was therefore a better indicator of palaeotemperatures in Westland.
This research has strengthened the New Zealand network of chronology sites, and confirmed that pink pine has great dendroclimatic value. The last 520 years of temperature fluctuations were reconstructed with a high degree of fidelity - the model developed in this thesis is currently the most accurate estimate of a temperature-growth relationship in the country.
