Mixed conifer, beech and hardwood forests are relatively common in Aotearoa/New
Zealand, but are not well studied. This thesis investigates the coexistence, regeneration
dynamics and disturbance history of a mixed species forest across an environmental
gradient of drainage and soil development in north Westland.
The aim was to investigate whether conifers, beech and non-beech hardwood species were
able to coexist on surfaces that differed in their underlying edaphic conditions, and if so to understand the mechanisms that influenced their regeneration on both poorly drained and
well drained soils. The site selected was an area of high tree species diversity on a lowland
0.8 km² post-glacial terrace at the base of Mount Harata in the Grey River Valley.
My approach was to use forest stand history reconstruction at two spatial scales: an
intensive within-plot study of stand dynamics (chapter 1) and a whole-landform approach
(chapter 2) that examined whether the dynamics identified at the smaller within-plot scale
reflected larger patterns across the terrace.
In chapter 1, three large permanent plots (0.3-0.7 ha) were placed at different points along
the drainage gradient, one plot situated in each of the mainly well-drained, poorly drained
and very poorly drained areas along the terrace. Information was gathered on species age
and size structures, spatial distributions of tree ages, species interactions, microsite
establishment preferences, patterns of stand mortality, and disturbance history in each plot.
There were differences in stand structure, composition and relative abundance of species
found between the well drained plot and the two poorer drained plots. On the well drained
site conifers were scarce, the beeches Nothofagus fusca and N. menziesii dominated the
canopy, and in the subcanopy the hardwood species Weinmannia racemosa and Quintinia
acutifolia were abundant. As drainage became progressively poorer, the conifers
Dacrydium cupressinum and Dacrycarpus dacrydioides became more abundant and
occupied the emergent tier over a beech canopy. The hardwoods W. racemosa and Q.
acutifolia became gradually less abundant in the subcanopy, whereas the hardwood
Elaeocarpus hookerianus became more so.
In the well drained plot, gap partitioning for light between beeches and hardwoods enabled
coexistence in response to a range of different sized openings resulting from disturbances
of different extent. In the two more poorly drained plots, species also coexisted by
partitioning microsite establishment sites according to drainage.
There were several distinct periods where synchronous establishment of different species
occurred in different plots, suggesting there were large disturbances: c. 100yrs, 190-200
yrs, 275-300 yrs and 375-425 yrs ago. Generally after the same disturbance, different
species regenerated in different plots reflecting the underlying drainage gradient. However,
at the same site after different disturbances, different sets of species regenerated,
suggesting the type and extent of disturbances and the conditions left behind influenced
species regeneration at some times but not others. The regeneration of some species (e.g.,
N. fusca in the well-drained plot, and Dacrydium in the poorer drained plots) was periodic
and appeared to be closely linked to these events. In the intervals between these
disturbances, less extensive disturbances resulted in the more frequent N. menziesii and
especially hardwood regeneration. The type of tree death caused by different disturbances
favoured different species, with dead standing tree death favouring the more shade-tolerant
N. menziesii and hardwoods, whereas uprooting created a mosaic of microsite conditions
and larger gap sizes that enabled Dacrycarpus, N. fusca and E. hookerianus to maintain
themselves in the poorly drained areas.
In chapter 2, 10 circular plots (c. 0.12 ha) were placed in well drained areas and 10
circular plots (c. 0.2 ha) in poorly drained plots to collect information on species
population structures and microsite preferences. The aims were to reconstruct species'
regeneration responses to a range of disturbances of different type and extent across the
whole terrace, and to examine whether there were important differences in the effects of
these disturbances.
At this landform scale, the composition and relative abundances of species across the
drainage gradient reflected those found in chapter 1. There were few scattered conifers in well drained areas, despite many potential regeneration opportunities created from a range
of different stand destroying and smaller scale disturbances.
Three of the four periods identified in chapter 1 reflected distinct terrace-wide periods of
regeneration 75-100 yrs, 200-275 yrs and 350-450 yrs ago, providing strong evidence of
periodic large, infrequent disturbances that occurred at intervals of 100-200 yrs. These
large, infrequent disturbances have had a substantial influence in determining forest
history, and have had long term effects on forest structure and successional processes.
Different large, infrequent disturbances had different effects across the terrace, with the
variability in conditions that resulted enabling different species to regenerate at different
times. For example, the regeneration of distinct even-aged Dacrydium cohorts in poorly
drained areas was linked to historical Alpine Fault earthquakes, but not to more recent
storms. The variation in the intensity of different large, infrequent disturbances at different
points along the environmental drainage gradient, was a key factor influencing the scale of
impacts. In effect, the underlying edaphic conditions influenced species composition along
the drainage gradient and disturbance history regulated the relative abundances of species.
The results presented here further emphasise the importance of large scale disturbances as a
mechanism that allows coexistence of different tree species in mixed forest, in particular
for the conifers Dacrydium, Dacrycarpus and the beech N. fusca, by creating much of the
environmental variation to which these species responded. This study adds to our
understanding of the effects of historical earthquakes in the relatively complex forests of
north Westland, and further illustrates their importance in the Westland forest landscape as
the major influential disturbance on forest pattern and history.
These results also further develop the 'two-component' model used to describe
conifer/angiosperm dynamics, by identifying qualitative differences in the impacts of
different large, infrequent disturbances across an environmental gradient that allowed for
coexistence of different species. In poorer drained areas, these forests may even be thought
of as 'three-component' systems with conifers, beeches and hardwoods exhibiting key
differences in their regeneration patterns after disturbances of different type and extent, and
in their microsite preferences.
A scanned copy of a photograph of the garden of Di Madgin's former home in the Red Zone, taken before the earthquakes. She describes the scene in the photograph as, "This is the courtyard that we made, to have an eating place at the back of the house. The tree in the neighbours' was a tree that Pete's brother stole on a school trip up in the mountains from a national park. They planted this red beech in the garden. It became the neighbourhood bird tree and the sound was fantastic in the evenings."
This documentary, made by TVNZ’s Natural History Unit (now NHNZ), charts the progress of the nor'west wind from its formation in the Tasman Sea across the Southern Alps to the Canterbury Plains and the east coast of the South Island. Along the way it dumps metres of precipitation on West Coast rain forest and snow on the Alps, then transforms to a dry, hot wind racing across the Plains. The film shows the wind's impact on the ecosystem and farming and muses on the mysterious effect it can have on humans. It screened as part of the beloved Wild South series.
Tree mortality is a fundamental process governing forest dynamics, but understanding tree mortality patterns is challenging
because large, long-term datasets are required. Describing size-specific mortality patterns can be especially difficult, due to
few trees in larger size classes. We used permanent plot data from Nothofagus solandri var. cliffortioides (mountain beech)
forest on the eastern slopes of the Southern Alps, New Zealand, where the fates of trees on 250 plots of 0.04 ha were
followed, to examine: (1) patterns of size-specific mortality over three consecutive periods spanning 30 years, each
characterised by different disturbance, and (2) the strength and direction of neighbourhood crowding effects on sizespecific
mortality rates. We found that the size-specific mortality function was U-shaped over the 30-year period as well as
within two shorter periods characterised by small-scale pinhole beetle and windthrow disturbance. During a third period,
characterised by earthquake disturbance, tree mortality was less size dependent. Small trees (,20 cm in diameter) were
more likely to die, in all three periods, if surrounded by a high basal area of larger neighbours, suggesting that sizeasymmetric
competition for light was a major cause of mortality. In contrast, large trees ($20 cm in diameter) were more
likely to die in the first period if they had few neighbours, indicating that positive crowding effects were sometimes
important for survival of large trees. Overall our results suggest that temporal variability in size-specific mortality patterns,
and positive interactions between large trees, may sometimes need to be incorporated into models of forest dynamics.
Documenting earthquake-induced ground deformation is significant to assess the characteristics of past and contemporary earthquakes and provide insight into seismic hazard. This study uses airborne light detection and ranging (LiDAR) and conducts multi-disciplinary field techniques to document the surface rupture morphology and evaluate the paleoseismicity and seismic hazard parameters of the Hurunui segment of the Hope Fault in the northern South Island of New Zealand. It also documents and evaluates seismically induced features and ground motion characteristics of the 2010 Darfield and 2011 Christchurch earthquakes in the Port Hills, south of Christchurch. These two studies are linked in that they investigate the near-field coseismic features of large (Mw ~7.1) earthquakes in New Zealand and produce data for evaluating seismic hazards of future earthquakes. In the northern South Island of New Zealand, the Australian-Pacific plate boundary is characterised by strike-slip deformation across the Marlborough Fault System (MFS). The ENE-striking Hope Fault (length: ~230 km) is the youngest and southernmost fault in the MFS, and the second fastest slipping fault in New Zealand. The Hope Fault is a major source of seismic hazard in New Zealand and has ruptured (in-part) historically in the Mw 7.1 1888 Amuri earthquake. In the west, the Hurunui segment of the Hope Fault is covered by beech forest. Hence, its seismic hazard parameters and paleoearthquake chronology were poorly constrained and it was unknown whether the 1888 earthquake ruptured this segment or not and if so, to what extent. Utilising LiDAR and field data, a 29 km-long section of the Hurunui segment of the Hope Fault is mapped. LiDAR-mapping clearly reveals the principal slip zone (PSZ) of the fault and a suite of previously unrecognised structures that form the fault deformation zone (FDZ). FDZ width measurements from 415 locations reveal a spatially-variable, active FDZ up to ~500 m wide with an average width of 200 m. Kinematic analysis of the fault structures shows that the Hurunui segment strikes between 070° and 075° and is optimally oriented for dextral strike-slip within the regional stress field. This implies that the wide FDZ observed is unlikely to result from large-scale fault mis-orientation with respect to regional stresses. The analysis of FDZ width indicates that it increases with increased hanging wall topography and increased topographic relief suggesting that along-strike topographic perturbations to fault geometry and stress states increase fault zone complexity and width. FDZ width also increases where the tips of adjacent PSZ strands locally vary in strike, and where the thickness of alluvial deposits overlying bedrock increases. LiDAR- and photogrammetrically-derived topographic mapping indicates that the boundary between the Hurunui and Hope River segments is characterised by a ~850-m-wide right stepover and a 9º-14° fault bend. Paleoseismic trenching at Hope Shelter site reveals that 6 earthquakes occurred at A.D. 1888, 1740-1840, 1479-1623, 819-1092, 439-551, and 373- 419. These rupture events have a mean recurrence interval of ~298 ± 88 yr and inter-event times ranging from 98 to 595 yrs. The variation in the inter-event times is explained by (1) coalescing rupture overlap from the adjacent Hope River segment on to the Hurunui segment at the study site, (2) temporal clustering of large earthquakes on the Hurunui segment, and/or (3) ‘missing’ rupture events. It appears that the first two options are more plausible to explain the earthquake chronologies and rupture behaviour on the Hurunui segment, given the detailed nature of the geologic and chronologic investigations. This study provides first evidence for coseismic multi-segment ruptures on the Hope Fault by identifying a rupture length of 44-70 km for the 1888 earthquake, which was not confined to the Hope River segment (primary source for the 1888 earthquake). LiDAR data is also used to identify and measure dextral displacements and scarp heights from the PSZ and structures within the FDZ along the Hurunui segment. Reconstruction of large dextrally-offset geomorphic features shows that the vertical component of slip accounts for only ~1% of the horizontal displacements and confirms that the fault is predominantly strike-slip. A strong correlation exists between the dextral displacements and elevations of geomorphic features suggesting the possibility of age correlation between the geomorphic features. A mean single event displacement (SED) of 3.6 ± 0.7 m is determined from interpretation of sets of dextral displacements of ≤ 25 m. Using the available surface age data and the cumulative dextral displacements from Matagouri Flat, McKenzie Fan, Macs Knob and Hope River sites, and the mean SED, a mean slip rate of 12.2 ± 2.4 mm/yr, and a mean recurrence interval of ~320 ± 120 yr, and a potential earthquake magnitude of Mw 7.2 are determined for the Hurunui segment. This study suggests that the fault slip rate has been constant over the last ~15000 yr. Strong ground motions from the 2010 Darfield (Canterbury) earthquake displaced boulders and caused ground damage on some ridge crests in the Port Hills. However, the 2011 Christchurch earthquake neither displaced boulders nor caused ground damage at the same ridge crests. Documentation of locations (~400 m a.s.l.), lateral displacements (8-970 cm), displacement direction (250° ± 20°) of displaced boulders, in addition to their hosting socket geometries (< 1 cm to 50 cm depth), the orientation of the ridges (000°-015°) indicate that boulders have been displaced in the direction of instrumentally recorded transient peak ground horizontal displacements nearby and that the seismic waves have been amplified at the study sites. The co-existence of displaced and non-displaced boulders at proximal sites suggests small-scale ground motion variability and/or varying boulder-ground dynamic interactions relating to shallow phenomena such as variability in soil depth, bedrock fracture density and/or microtopography on the bedrock-soil interface. Shorter shaking duration of the 2011 Christchurch event, differing frequency contents and different source characteristics were all factors that may have contributed to generating circumstances less favourable to boulder displacement in this earthquake. Investigating seismically induced features, fault behaviour, site effects on the rupture behaviour, and site response to the seismic waves provides insights into fault rupture hazards.
