Tree mortality is a fundamental process governing forest dynamics, but understanding tree mortality patterns is challenging
because large, long-term datasets are required. Describing size-specific mortality patterns can be especially difficult, due to
few trees in larger size classes. We used permanent plot data from Nothofagus solandri var. cliffortioides (mountain beech)
forest on the eastern slopes of the Southern Alps, New Zealand, where the fates of trees on 250 plots of 0.04 ha were
followed, to examine: (1) patterns of size-specific mortality over three consecutive periods spanning 30 years, each
characterised by different disturbance, and (2) the strength and direction of neighbourhood crowding effects on sizespecific
mortality rates. We found that the size-specific mortality function was U-shaped over the 30-year period as well as
within two shorter periods characterised by small-scale pinhole beetle and windthrow disturbance. During a third period,
characterised by earthquake disturbance, tree mortality was less size dependent. Small trees (,20 cm in diameter) were
more likely to die, in all three periods, if surrounded by a high basal area of larger neighbours, suggesting that sizeasymmetric
competition for light was a major cause of mortality. In contrast, large trees ($20 cm in diameter) were more
likely to die in the first period if they had few neighbours, indicating that positive crowding effects were sometimes
important for survival of large trees. Overall our results suggest that temporal variability in size-specific mortality patterns,
and positive interactions between large trees, may sometimes need to be incorporated into models of forest dynamics.
We examined the stratigraphy of alluvial fans formed at the steep range front of the Southern Alps at Te Taho, on the north bank of the Whataroa River in central West Coast, South Island, New Zealand. The range front coincides with the Alpine Fault, an Australian-Pacific plate boundary fault, which produces regular earthquakes. Our study of range front fans revealed aggradation at 100- to 300-year intervals. Radiocarbon ages and soil residence times (SRTs) estimated by a quantitative profile development index allowed us to elucidate the characteristics of four episodes of aggradation since 1000 CE. We postulate a repeating mode of fan behaviour (fan response cycle [FRC]) linked to earthquake cycles via earthquake-triggered landslides. FRCs are characterised by short response time (aggradation followed by incision) and a long phase when channels are entrenched and fan surfaces are stable (persistence time). Currently, the Te Taho and Whataroa River fans are in the latter phase. The four episodes of fan building we determined from an OxCal sequence model correlate to Alpine Fault earthquakes (or other subsidiary events) and support prior landscape evolution studies indicating ≥M7.5 earthquakes as the main driver of episodic sedimentation. Our findings are consistent with other historic non-earthquake events on the West Coast but indicate faster responses than other earthquake sites in New Zealand and elsewhere where rainfall and stream gradients (the basis for stream power) are lower. Judging from the thickness of fan deposits and the short response times, we conclude that pastoral farming (current land-use) on the fans and probably across much of the Whataroa River fan would be impossible for several decades after a major earthquake. The sustainability of regional tourism and agriculture is at risk, more so because of the vulnerability of the single through road in the region (State Highway 6).
